A flower touched by the witch who once dreamt of burning away all the demons in the world. The anonymous flames affectionately caress the hands of those who touch it.
She was then recruited into the Fatui by Pierro. She was given a Cryo delusion to suppress her flames, and became known as La Signora, the Eighth of the Eleven Fatui Harbingers.
Signora, also known by her alias "The Fair Lady" and her real name Rosalyne-Kruzchka Lohefalter, was the 8th of the Eleven Harbingers of the Fatui.
She can be seen wielding both Cryo and Pyro elements. She uses a Cryo Delusion to suppress her natural affinity toward using the art of liquid flame. The art of liquid flame has been long lost to history, and only Signora knows its intricacies.
She is first seen at the end of Act 3 of the Prologue, after the Traveler has assisted Dvalin and returned the Holy Lyre der Himmel.
She appears with Fatui agents and mages, catching the Traveler and Venti off guard as they exit the Mondstadt Cathedral. La Signora makes quick work of the Traveler, Paimon, and Venti, apprehending them with a single snap that freezes Paimon entirely and blows her away, and freezes Venti's legs in place so he cannot escape. Fatui agents hold back the Traveler as she taunts Venti. When Venti returns the favor, she becomes enraged and plunges her arm into his chest, taking his Gnosis. She leaves him to fall on the ground after kicking him once more, and leaves the Traveler and Paimon behind before any Knights of Favonius show up and catch her in the act.
Signora reappears in Act 3 of Archon Quest Chapter I, where she is seen with Childe and Zhongli, the latter of which forfeiting his Gnosis to her as per terms of his contract with the Tsaritsa. Signora calls Zhongli's speech during the affair "sanctimonious" and taunts Childe and the Traveler, before leaving alone to report back to the Tsaritsa.
Signora makes her last living appearance Act 3 of Chapter II, where she meets with the Raiden Shogun within Tenshukaku. Here, she flaunts her status as a Snezhnayan diplomat, as well as revealing she had incapacitated Kujou Sara before the Traveler's arrival. The Traveler realizes that Signora is responsible for all of the Fatui's plots on Inazuma, including the Vision Hunt Decree and the distribution of Delusions to members of the Sangonomiya Resistance, and challenges Signora to a duel before the throne. During the duel, Signora then reveals her true original power. After being defeated by the Traveler, she reverts from her Crimson Witch form, considerably weakened from the fight. As per the rules of the duel, Signora is formally executed by the Raiden Shogun. In her final moments, Signora threatens the Raiden Shogun, calling her and the Traveler "filthy rats," before she is slain by the Shogun's Musou no Hitotachi. Signora disintegrates into ashes.
Signora acts very posh and haughty on her introduction, calling Barbatos a "rodent ruler", although given her position as a Harbinger, she may already know that he was the weakest of The Seven to begin with. She also takes great offense when he suggests that she's only powerful because of the Tsaritsa, and brutally takes his Gnosis immediately after — though whether her anger stems from the insult to her archon or the insult to her own power is unknown.
However, she is quite different when interacting with Zhongli, on account of his formal contract with Tsaritsa. While she mocks the Traveler over the incident in Mondstadt and belittles fellow Harbinger Childe, she is much more professional with Zhongli himself. Additionally, she was able to keep the contract confidential, even keeping Childe's "eavesdropping ears" from gleaning such information, to which Zhongli praises.
Signora seems to hold the sentiment that only the strongest can survive, as several mention that she is very strong herself and hold faith in her skills, while Childe states that she uses lethal force against her recruits.